IOWA CITY, Iowa (AP) -- There's no question that Michigan State's defense is among the best in the nation.
If the Spartans throw the ball and play special teams like they did against Iowa, they'll be a serious contender in the Big Ten this season.
Connor Cook threw for a career-high 277 yards and two touchdowns and Michigan State opened conference play with a 26-14 win over the Hawkeyes on Saturday.
Darqueze Dennard had a pair of interceptions and freshman Michael Geiger had four field goals for the Spartans (4-1, 1-0 Big Ten), who held the Hawkeyes (4-2, 1-1) scoreless in the second half.
"We were tough to run the ball against, may be an understatement. We came up with two big turnovers at the right times," Michigan State coach Mark Dantonio said.
Mark Weisman ran for just nine yards on seven carries for the Hawkeyes. They finished with just 23 yards rushing -- or two less than Michigan State punter Mike Sadler had on one fake -- and lost for the first time since its season opener.
Iowa, which averaged over 50 rushes a game entering play, was forced to throw the ball 46 times against the Spartans.
"To try to move the ball effectively on them, you've got to be mixed and balanced. It was tough sledding out here," Iowa coach Kirk Ferentz said. "We were hoping we could run it effectively. But the way they're built, they make it very difficult to do that."
Geiger and Sadler teamed up to headline a huge day for Michigan State's special teams.
Up 6 early in the fourth quarter, Sadler took off on a run that nullified a big stand for Iowa's defense. The successful fake set up a 49-yard field goal by Geiger that put the Spartans up 23-14 with 13:28 left, which proved to be insurmountable against the Spartans.
"It's all about timing and execution. We just felt like we had the time and that it was the right moment to do it. I just wanted our players to know that they're aggressive on the football field and sometimes coaches need to take a risk too," Dantonio said.
Jake Rudock threw for 241 yards and two touchdowns for the Hawkeyes. But Dennard picked him off twice, including an interception with 5:09 left that largely ended the suspense.
Michigan State hasn't allowed more than 17 points in five games this season.
"When you have a defense that can hold a powerful offense like Iowa to under 30 yards rushing, that's incredible," Cook said. "We go into every single football game and think, we score three touchdowns and we're going to win. We didn't score three touchdowns but we had a lot of field goals."
Michigan State's stellar second half made it easy to forget how much momentum Iowa had at halftime.
The Hawkeyes found some life in their passing game and quickly turned that 10-point deficit into a 14-10 lead at the break.
Rudock completed his last 11 passes of the first half for 138 yards and two touchdowns -- by far the best stretch of his young career.
Rudock beat a Michigan State blitz by finding Damon Bullock in the flat for a 47-yard touchdown, and he hit C.J. Fiedorowicz for a 10-yard touchdown pass with 1:10 left in the second quarter.
That would be it for the Hawkeyes. They played most of the game without leading receiver Kevonte Martin-Manley, who went out with a leg injury that Ferentz said shouldn't be a lingering one.
"They've got the momentum; we've got to get it back," Dantonio said. "The defense helped out. They usually do."
NO. 1 ALABAMA 45, GEORGIA ST. 3
TUSCALOOSA, Ala. -- AJ McCarron completed 15 of 16 passes for 166 yards and four touchdowns in the first half and No. 1 Alabama rolled to a victory over Georgia State.
The Crimson Tide (5-0) jumped ahead 38-0 by halftime against the Panthers (0-5), a first-year Football Bowl Subdivision team.
McCarron led Alabama to touchdowns on each of his five possessions before leaving the game. He completed his first 12 passes, putting him in a three-way tie for the Tide's third-longest streak.
The four touchdown passes tied his career high and was the seventh time McCarron has reached that number. Alabama outgained the Panthers 477-175 and held them to 15 yards rushing.
The Tide hasn't allowed a touchdown in the past three games, giving up only three field goals.
Georgia State ended Alabama's shutout bid with Wil Lutz's school-record 53-yard field goal to cap the second half's opening drive.
NO. 2 OREGON 57, COLORADO 16
BOULDER, Colo. -- Marcus Mariota threw five touchdown passes and ran for two scores as No. 2 Oregon brushed off some early trickery to rout Colorado.
The Ducks' sophomore connected for two touchdowns each with Josh Huff and Bralon Addison, and played only 2 1/2 quarters.
The Buffaloes had Folsom Field in a frenzy when they kept things tight early on, but the crowd didn't like it when Colorado kept going for field goals while the Ducks were piling up the points.
Oregon (5-0, 2-0 Pac-12) has scored at least 55 points in all of its games under first-year coach Mark Helfrich, a former offensive coordinator at Colorado, who took over when Chip Kelly left to coach the Philadelphia Eagles.
Forcing a three-and-out after the opening onside kick, a 75-yard TD toss from receiver Paul Richardson on a double-reverse and a fourth-down stop had the 40-point-underdog Buffaloes (2-2, 0-2) up 10-8 early.
NO. 3 CLEMSON 49, SYRACUSE 14
SYRACUSE, N.Y. -- Tajh Boyd kept his Heisman Trophy aspirations intact, throwing for 455 yards and five touchdowns in three quarters of work, and No. 3 Clemson beat Syracuse to spoil the Orange's Atlantic Coast Conference debut.
Boyd, who matched his school record in TD passes, hit Adam Humphries with scoring passes of 60 and 42 yards in the first quarter to help stake the Tigers to a big early lead, and they held the Orange at bay.
Clemson (5-0, 3-0 ACC) entered the game as one of just 20 undefeated teams remaining in the Bowl Subdivision, and the Tigers made sure they wouldn't slip up against the Orange (2-3, 0-1) and their raucous Homecoming crowd of 48,961 by taking a 21-0 first-quarter lead.
NO. 6 GEORGIA 34, TENN. 31, OT
KNOXVILLE, Tenn. -- Marshall Morgan kicked a 42-yard field goal in overtime after No. 6 Georgia scored a tying touchdown with 5 seconds left in regulation as the Bulldogs overcame numerous injuries to beat Tennessee.
Tennessee's Alton "Pig" Howard capped the first overtime possession by losing control of the ball as he dove toward the front corner of the end zone. Howard's play was initially ruled a 7-yard touchdown, but replays showed he fumbled prior to crossing the goal line, turning the potential score into a turnover and touchback.
Georgia (4-1, 3-0 SEC) forced overtime on Aaron Murray's 2-yard touchdown pass to Rantavious Wooten to make it 31-all. Rajion Neal's 7-yard run had put Tennessee (3-2, 0-1) ahead 31-24 with 1:54 left.
Georgia running back Keith Marshall injured his right knee in the first quarter. Receivers Michael Bennett and Justin Scott-Wesley and punter Colin Barber also were injured.
NO. 7 LOUISVILLE 30, TEMPLE 7
PHILADELPHIA -- Teddy Bridgewater threw for 348 yards and two touchdowns to keep No. 7 Louisville undefeated with a win over winless Temple.
Bridgewater made it look easy against the overwhelmed Owls (0-5, 0-2 American Athletic Conference). He threw for 228 yards in the first half to help the Cardinals roll to a 24-0 lead.
The Cardinals (5-0, 1-0) lost a bid for their second straight shutout when Temple scored with 38 seconds left.
NO. 8 FSU 63, NO. 25 MARYLAND 0
TALLAHASSEE, Fla. -- Eighth-ranked Florida State stayed undefeated in Atlantic Coast Conference play with a victory over No. 25 Maryland.
The Seminoles (5-0, 3-0) rode the arm of quarterback Jameis Winston, who completed 23-of-32 passes for 393 yards and five touchdowns.
Maryland (4-1, 0-1) never got anything going and was lost after quarterback C.J. Brown left the game with an injury in the second quarter. The No. 2 total offense in the ACC was held to 234 yards.
NO. 11 OKLAHOMA 20, TCU 17
NORMAN, Okla. -- Brennan Clay broke a 76-yard touchdown run late in the fourth quarter and No. 11 Oklahoma held off TCU 20-17 on Saturday night.
Trey Millard had a touchdown run for the Sooners (5-0, 2-0 Big 12) and Michael Hunnicutt hit field goals of 39 and 32 yards. Oklahoma and Texas will meet next Saturday in the annual Red River Rivalry in Dallas with first place in the conference on the line.
Oklahoma has won 10 straight Big 12 games, its third-longest streak in conference play since the league formed in 1996. The Sooners have won their last three against TCU (2-3, 0-2) since the Horned Frogs won in Norman in 2005.
NO. 14 MIAMI 45, GEORGIA TECH 30
MIAMI GARDENS, Fla. -- Stephen Morris threw for three touchdowns, Duke Johnson rushed for 184 yards and No. 14 Miami remained unbeaten after shaking off a problematic opening quarter to beat Georgia Tech.
Phillip Dorsett, Clive Walford and Allen Hurns caught scoring passes for Miami (5-0, 1-0 Atlantic Coast Conference), which scored 21 points in the final 7:15. Dallas Crawford ran for two scores in the fourth quarter, and Ladarius Gunter added a 30-yard interception return for another touchdown with 1:08 left.
NO. 20 TEXAS TECH 54, KANSAS 16
LAWRENCE, Kan. -- Freshman sensation Baker Mayfield passed for 368 yards but was helped off the field with an apparent leg injury in the third quarter of No. 20 Texas Tech's victory over Kansas.
After falling behind in the first quarter 10-0, the unbeaten Red Raiders reeled off 54 straight points, going to 5-0 (2-0 Big 12) under first-year head coach Kliff Kingsbury for the first time since 2008.
NO. 21 OKLA. ST. 33, KANSAS 29
STILLWATER, Okla. -- J.W. Walsh found Charlie Moore for a 6-yard touchdown pass with 4:16 left in the fourth quarter to help No. 21 Oklahoma State rally for a win over Kansas State.
After falling behind 29-23 in the fourth quarter, Oklahoma State (4-1, 1-1 Big 12 Conference) drove 75 yards in six plays. Walsh was 4-of-5 passing on the drive, including the go-ahead touchdown pass to Moore.
NO. 23 FRESNO ST. 61, IDAHO 14
MOSCOW, Idaho -- Derek Carr threw five touchdown passes as No. 23 Fresno State pounded Idaho 61-14 on Saturday, scoring at least 40 points for the fifth time this season.
Davante Adams caught three touchdown passes for Fresno State (5-0), which is off to its best start since 2001, when the Bulldogs won their first six games.
Idaho (1-5), coming off a win over Temple in coach Paul Petrino's first season, didn't score until the fourth quarter against the first ranked opponent to play in the Kibbie Dome since Boise State in 2010.
NEBRASKA 39, ILLINOIS 19
LINCOLN, Neb. -- Ameer Abdullah ran for a career-high 225 yards and two touchdowns and Nebraska's maligned defense was much improved in a victory over Illinois.
Tommy Armstrong, starting his second straight game in place of injured quarterback Taylor Martinez, led the Cornhuskers (4-1, 1-0 Big Ten) to touchdowns on his first three series and was solid throughout.
Armstrong was 8 of 13 for 135 yards.
Just as big as Armstrong's performance was that of Nebraska's defense, which had been one of the nation's worst through four games.
Illinois (3-2, 0-1), which came in averaging 40 points a game, didn't score its first touchdown until the middle of the third quarter. Twice when the Illini looked as if they would make a game of it, the Huskers made big stops.
Josh Ferguson led the Illini with 114 yards on 19 carries and another 82 yards on eight catches.
INDIANA 44, PENN STATE 24
BLOOMINGTON, Ind. -- Nate Sudfeld threw two touchdown passes and Tre Roberson ran for two more scores, leading Indiana to its first win ever against Penn State.
The Hoosiers (3-2, 1-0 Big Ten) capped their five-game home stand by ending a 16-game losing streak to the Nittany Lions (3-2, 0-1).
It looked like Penn State might keep the streak alive after taking its first lead of the game on a 26-yard TD pass from Christian Hackenberg to Allen Robinson early in the third quarter.
But Indiana answered with a scoring flurry.
Tevin Coleman scored on a 44-yard run and Roberson scored on a 2-point conversion run. Roberson then added a 2-yard TD run early in the fourth and after Penn State failed on fourth-and-2 from its own 33, Kofi Hughes made a catch for a 36-yard TD and a 35-17 lead. Roberson followed with a 9-yard TD run to seal it.
BUFFALO 42, E. MICHIGAN 14
BUFFALO, N.Y. -- Branden Oliver gained 150 yards rushing and scored twice as Buffalo cruised to a victory over Eastern Michigan.
The Bulls (3-2, 1-0 Mid-American Conference) broke the 40-point mark in back-to-back contests for the first time since 1981 and snapped a five-game losing streak against the Eagles (1-4, 0-2 MAC).
Oliver set the tone early with a 60-yard touchdown run on Buffalo's first possession of the game before adding a 2-yard touchdown run later in the first quarter.
With the Bulls leading 21-0, Eastern Michigan got on the board with a second-quarter touchdown but Buffalo's Devin Campbell returned the ensuing kickoff 96 yards for a touchdown.
Campbell also added a 1-yard touchdown run in the third quarter before backups for both schools took reps in the fourth quarter with Buffalo up 42-7.
Ryan Brumfield had a 75-yard touchdown run late in the fourth quarter for the Eagles.
BALL STATE 48, VIRGINIA 27
CHARLOTTESVILLE, Va. -- Keith Wenning threw for two touchdowns and ran for another as Ball State defeated Virginia.
Jahwan Edwards added three scoring runs for Ball State (5-1), which shredded a Virginia (2-3) defense that came in allowing only 299 yards per game. The Cardinals finished with 506 yards.
Wenning had TD passes of 11 yards to Willie Snead in the second quarter and 72 yards to Jordan Williams -- a fourth-quarter back-breaker that put the Cardinals up 41-27 and sent many in the crowd of 38,228 scurrying for the exits.
Edwards' 17-yard TD run sealed it. He also had scoring runs of 3 and 32 yards.
Virginia's David Watford was 21 of 36 for 209 yards and one interception. Kevin Parks ran for two touchdowns for Virginia. The Cavaliers looked explosive at times as they compiled 459 yards, but they were done in by missed opportunities, turnovers and penalties.
C. MICHIGAN 21, MIAMI (OH) 9
OXFORD -- Saylor Lavallii rushed for 151 yards and two touchdowns to lead Central Michigan to a victory over Miami (Ohio).
Lavallii rushed 19 yards for a score to give the Chippewas (2-4, 1-1 Middle Atlantic Conference) a 7-3 lead early in the second quarter. Miami (0-5, 0-1) responded in the third quarter when Austin Boucher found David Frazier for the 10-yard score.
Central Michigan took the next drive and marched on a 16-play, 84-yard scoring drive, capped off with Lavallii's 3-yard score. Maurice Shoemaker-Gilmore added a 16-yard scoring run with 6:13 left to play, and the Chippewas let their defense do the rest.
Miami's Austin Gearing finished with 108 yards on 15 carries but was unable to find the end zone. This is the third time this season that Miami was held to less than 10 points.
Central Michigan will take on Ohio next week while Miami will play Massachusetts.
OHIO 43, AKRON 3
AKRON -- Tyler Tettleton threw for two touchdowns as Ohio opened Mid-American Conference play with a victory over Akron.
Ohio (4-1, 1-0 Mid-American) started slow but scored 21 unanswered points in the second quarter to take a 30-3 halftime lead.
Tettleton finished the game with 171 total yards on 14-of-21 passing. Donte Foster was his top receiver as he pulled in four passes for 76 yards and one touchdown.
Kyle Pohl lead Akron (1-5, 0-2), throwing for 101 yards. Robert Stein's 31-yard field goal was Akron's only score.
The Bobcat defense stymied the Zips all day, limiting Akron to 157 total yards, and sacking Pohl eight times for total losses of 45 yards. Antwa Crutcher and Ty Branz combined for 13 tackles and 3 sacks to lead the Ohio defense. Tarell Basham also had two sacks.
TOLEDO 47, W. MICHIGAN 20
TOLEDO -- David Fluellen rushed for 220 yards and four touchdowns as Toledo routed Western Michigan 47-20 in a Mid-American Conference matchup on Saturday.
Toledo (3-3, 2-1) ran up 360 rushing yards and limited Western Michigan (0-6, 0-2) to 87 yards on the ground. Toledo also won the turnover battle, as the Broncos lost two fumbles and threw two interceptions.
The Rockets took off in the first quarter, scoring on their first two possessions. Following an interception by Chaz Whittaker, Terrance Owens hit Bernard Reedy for a 40-yard touchdown strike on Toledo's first offensive play of the game. Their second drive culminated with Fluellen's 7-yard TD scamper.
Fluellen added three more scores on runs of 6, 66 and 2 yards, and Toledo led by 40 at the end of the third quarter.
Western Michigan's Andrew Haldeman kicked two field goals and Dareyon Chance pounded in a 1-yard touchdown run in the fourth.
BOWLING GREEN 28, UMASS 7
BOWLING GREEN -- Matt Johnson passed for 167 yards and three touchdowns to lead Bowling Green to a 28-7 homecoming win over the University of Massachusetts in a Mid-Atlantic Coast matchup Saturday.
Johnson spread the attack, sending touchdown throws to three different receivers, including a 51-yard pass to Ronnie Moore in the first quarter to give the Falcons (5-1, 3-0) a lead they'd keep over the winless Minutemen (0-5, 0-1). Tyler Beck caught a 23-yard pass for a touchdown in the third quarter, followed by a 9-yard TD catch by Heath Jackson five minutes later. William Houston also punched in a 1-yard touchdown run.
The only UMass score came late in the fourth quarter on a 24-yard touchdown pass from Mike Wegzyn to Tajae Sharpe.
Bowling Green also got another big game from Travis Greene, who was kept out of the end zone but rushed for 160 yards.
N. ILLINOIS 38, KENT ST. 24
KENT -- Cameron Stingily ran for a career-high 266 yards and two scores as Northern Illinois remained undefeated with a win over Kent State.
Northern Illinois (5-0, 1-0 Mid-American) trailed 24-21 in the third quarter but proceeded to score 17 unanswered points, beginning with a 22-yard touchdown run by James Spencer midway through the quarter. Spencer then scored on an 11-yard touchdown pass from Jordan Lynch on the Huskies' following possession. Northern Illinois finished with 698 yards of total offense, 454 yards rushing.
Kent State (2-4, 1-2) took the lead early in the third with a 28-yard touchdown pass from Colin Reardon to Tyshon Goode but failed to score after that.